Jouelle Persians

**Patterns Produced by the White Piebald Spotting Gene**
All Black = Grade 1 Mitted (Ss)= Grades 2 to 3, low grade white Bicolor (Ss) = Grades 4 to 5, medium grade white Harlequin (SS) = Grades 6 to 8, high grade white Van (SS) = Grade 9, highest grade white All White = Grade 10

Variations in Expression of the Piebald Spotting Gene
SIRE	DAM	KITTENS	Description of expression in kittens:
Mitted (Ss)	Mitted (Ss)	Mitted Bicolor	Grades 1 to 5: Solid Low grade white (mitted pattern) Medium grade white (bicolor)
Mitted (Ss)	Bicolor (Ss)	Mitted Bicolor	Grades 2 to 5: Low grade white (mitted pattern) Medium grade white (bicolor)
Bicolor (Ss)	Bicolor (Ss)	Mitted Bicolor	Grades 2 to 5: Low grade white (mitted pattern) Medium grade white (bicolor)
Harlequin (SS)	Mitted (Ss)	Mitted Bicolor Harlequin Van	Grades 2 to 9 Low grade white (mitted pattern) Medium grade white (bicolor) High grade white (Harlequin) Highest grade white (Van)
Harlequin (SS)	Bicolor (Ss)	Mitted Bicolor Harlequin Van	Grades 2 to 9 Low grade white (mitted pattern) Medium grade white (bicolor) High grade white (Harlequin) Highest grade white (Van)
Van (SS)	Mitted (Ss)	Mitted Bicolor Harlequin Van	Grades 2 to 9 Low grade white (mitted pattern) Medium grade white (bicolor) High grade white (Harlequin) Highest grade white (Van)
Van (SS)	Bicolor (Ss)	Mitted Bicolor Harlequin Van	Grades 2 to 9 Low grade white (mitted pattern) Medium grade white (bicolor) High grade white (Harlequin) Highest grade white (Van)
Van (SS)	Van (SS)	Harlequin Van	Grades 6 to 9 High grade white (Harlequin) Highest grade white (Van)

Cats have only two basic colors (Black and Red). All other colors are variations produced by the affect of the dilute (recessive) and/or agouti factors in the coloring matter in the hair fibers (melanin).

Male cats carry only one color (Black or Red)
Female cats carry two colors (Black and Black); (Black and Red); or (Red and Red).
Male kittens get their color from their dam (i.e..Black dam produces Black sons).
Female kittens get one color from each parent (i.e..Black dam x Red sire produces Tortoiseshell daughters).
If both parents are dilute (Blue, Cream), then the color of all offspring must always be dilute.

White is not a color. It is dominant to all other colors and when present masks (covers over) the cat's genetic color.

A White cat must have a white parent.

Piebald/White Spotting represents areas of skin in which the pigment-producing cells are absent. This effect is caused by a dominant gene S, with variable expression. The amount of white can vary from a few small patches to large white areas covering most of the body depending partly on whether the cat possesses one or two of the genes. Moderate white spotting usually affects the under-parts of the body. Bi-Colors may not be true breeding because gene S is incompletely dominant. One parent must be a Bi-Color to produce Bi-Color kittens.

The piebald spotting gene is inherited as an incomplete dominant gene. This is what causes the variation in penetrence and expressibility. EXPRESSIVITY - Refers to the degree of phenotypic expression of a gene or genotype. Absence of expressivity is also called nonpenetrance.

Vans (Harlequins) are produced from a (Bi-Color x Bi-Color), and (Bi-Color x Van Bi-Color). When both sire and dam are Vans, then all kittens produced will be Van Bi-Colors. A Van is a white cat with color confined to the extremities; head, tail, and legs, one or two small body spots or color patches on the body are allowable.

When a Solid color is bred to a Bi-Color (or Calico), you would expect 50% of the offspring to be a Solid color and 50% of the offspring to be Bi-Color and Calico.
When a Bi-Color is bred to a Bi-Color (or Calico), you would expect 25% of the offspring to be a Solid color, 50% of the offspring to be Bi-Color, and 25% of the offspring to be Van Bi-Colors.
When a Bi-Color (or Calico) is bred to a Van Bi-Color, you would expect 50% of the offspring to be Bi-Color and 50% of the offspring to be Van Bi-Color.
When Van Bi-Color is bred to a Van Bi-Color, only Van Bi-Color offspring can produced.
When a Solid color is bred to a Van Bi-Color all resulting offspring will be Bi-Color or Calico; Van Bi-Color kittens can not be produced when one parent is a Solid color.